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Brown algae

Brown algae or Phaeophyta are mainly found in our area in the intertidal zone and in shallow water. They usually have a characteristic brownish colour. Brown algae include most of the more or less leaf-shaped seaweeds that often form huge forests in the oceans. In the Mediterranean these large seaweeds are less common.

Here you can skip the introduction and go directly to the species.

Brown algae
typical community of brown algae

Brown algae
Brown algae occur especially on slate and granite; on marble they are less common.

Brown algae
They often form dense populations on rocks both in the intertidal zone and in the shallow water.

Most brown algae around Naxos show a shrub-like growth form. They form extensive colonies, especially on slate in shallow water. The most common genus is Cystoseira with numerous, partly endemic species, which are generally difficult to distinguish. Apart from Cystoseira, Dictyota dichotoma and Peacock’s tail (Padina pavonica) are the most common species.



Brown algae on Naxos

The Photo gallery of the marine plants gives an overview over the species.

A note on identification: Some species of algae can be identified easily and reliably. In many cases, however, there are several related species that are difficult to distinguish. Often, examination under a microscope is necessary for correct identification. Another problem with the identification is that not all species are included in the field guides; even using the internet, it is difficult to find information on many species. Often, the species identified with the help of books or photos on the internet look quite different from our specimen here. Accordingly, many of the identifications are unfortunately quite uncertain, and some species must remain completely unidentified.

Here you can jump directly to the species (return with the back arrow or by swiping back): Dictyota  –  Taonia atomaria  –  Zonaria tournefortii  –  Padina pavonica  –  Stypopodium zonale  –  Cystoseira  –  Sporochnus pedunculatus

Dictyota dichotoma, J. V. Lamouroux

common in shallow water, especially on slate

Dictyota dichotoma
Dictyota dichotoma is easily recognisable by its ribbon-shaped, forked thallus.

Dictyota dichotoma
The fine dots on the thallus are the spore-producing areas.

Dictyota dichotoma
Here, the spore-producing areas are arranged in indistinct transverse lines.

Dictyota dichotoma
Sometimes Dictyota dichotoma shows a greenish iridescence.

Dictyota dichotoma var. intericata, Greville

much rarer, found on granite in the south-west of Naxos; at the waterline

Dictyota dichotoma var. intericata
Dictyota dichotoma var. intericata has much finer thallus sections, which are approximately the same width along their entire length.

Dictyota fasciola, J. V. Lamouroux

very rare, perhaps in deeper water

Dictyota fasciola
In this species, the narrow thallus sections become thinner towards the ends; the plant forms an upward-spreading tuft that is attached to the substrate with an adhesive organ.

Taonia atomaria, J. Agardh

rare, in slightly deeper water

Taonia atomaria
This species is closely related to the genus Dictyota, but it does not occur directly at coast; occasionally we find it washed ashore.

Taonia atomaria
Its very thin thallus is divided into lobes or narrow bands that taper towards the ends. The spores are produced in the darker transverse bands.

Zonaria tournefortii, Montagne

very rare

Zonaria tournefortii
The thallus of this species is irregularly divided into larger and smaller sections, which always have a broad, rounded end. It shows clearly visible transverse bands.

Peacock’s tail, Padina pavonica, L.

common in shallow water, especially on slate

Padina pavonica

Padina pavonica
The unmistakable Peacock’s tail is one of the common brown algae in our region.

Stypopodium zonale ?, Papenfuss

rare, inconspicuous and small

Stypopodium zonale?
This very small alga forms a thin, fan-shaped thallus. It shows a very strong blue-green iridescence.

Stypopodium zonale?
Here you can see the fine transverse bands on the thallus.

Cystoseira

very common in the infralittoral zone from the waterline downwards

Cystoseira
The genus Cystoseira is particularly typical of the Mediterranean region. Most species are difficult to tell apart.

Cystoseira
The species of Cystoseira have round stems, from which many branched fronds emerge, often bearing small pneumatocysts (air filled swim bodies) or spines.

Cystoseira
Some species are finely branched, such as the one pictured here.

Identification of the species is not easy. Here an attempt:

Cystoseira
Cystoseira abies-marina has no pneumatocysts, but sawtooth-like spines.

Cystoseira
Cystoseira barbata has no spines, but slightly inflated pneumatocysts arranged in chains.

Cystoseira
Cystoseira mediterranea (middle) shows a greenish iridescence.

Cystoseira
This species is supposed to have small thorns clustered densely at the ends of the branches; however, these are missing here and the branching is quite sparse, making identification uncertain.

Cystoseira
Here is a specimen with finer branches; possibly a different species.

Cystoseira compressa, Gerloff & Nizamuddin

Unlike other species, Cystoseira compressa occurs primarily in the eulittoral zone, where it is very common.

Cystoseira compressa
Cystoseira compressa differs from other species in its flat and branched thalli. It has neither pneumatocysts nor spines.

Cystoseira compressa

Cystoseira compressa
Numerous white dots (spore-forming zones?) sit on the thallus.

unidentified species

rare, on marble in the eulittoral


This stiff alga with short, strong, ribbon-shaped leaves is rather rare in our region. It occurs close to the waterline on marble.


At first glance, the species resembles a Fucus, but it grows upright, is very stiff and has only an indistinct, broad midrib; the pneumatocysts typical of this genus are also missing.


The smooth leaves are covered with many small ‘bubbles’ which probably produce the spores.

Sporochnus pedunculatus ?, C. Agardh

in deeper water, often washed ashore at the beach

Sporochnus pedunculatus?
This species is particularly delicate and fine-threaded; when floating in water, it takes on a somewhat spherical shape.

Sporochnus pedunculatus?
Its structure resembles that of the red algae Wrangelia with its tufted side branches, but it is significantly more delicate.

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